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Committee on Evolutionary Biology

Department Website:


  • Michael Coates

Associate Chair

  • Shannon Hackett


  • Kenneth Angielczyk, Field Museum
  • John Bates, Field Museum
  • Joy Bergelson, Ecology and Evolution
  • Rüdiger Bieler, Field Museum
  • Michael Coates, Organismal Biology and Anatomy
  • Maureen Coleman, Geophysical Sciences
  • Katherine Cronin, Lincoln Park Zoo
  • Martin Feder, Organismal Biology and Anatomy
  • Michael J. Foote, Geophysical Sciences
  • Lance Grande, Field Museum
  • Shannon Hackett, Field Museum
  • Lawrence Heaney, Field Museum
  • Patrick Herendeen, Chicago Botanic Garden
  • Andrew Hipp, Morton Arboretum/Herbarium
  • Robert Ho, Organismal Biology and Anatomy
  • David Jablonski, Geophysical Sciences
  • Susan M. Kidwell, Geophysical Sciences
  • Marcus Kronforst, Ecology and Evolution
  • Robert Lacy, Brookfield Zoo
  • Scott Lidgard, Field Museum
  • Sarah London, Psychology
  • Manyuan Long, Ecology and Evolution
  • Thorston Lumbsch, Field Museum
  • Zhe-Xi Luo, Organismal Biology and Anatomy
  • Peter Makovicky, Field Museum
  • Robert D. Martin, Field Museum
  • Jill Mateo, Comparative Human Development
  • Lance Miller, Chicago Zoological Society (Brookfield Zoo)
  • R. Michael Miller, Argonne National Laboratory
  • Gregory M. Mueller, Chicago Botanic Garden
  • Salikoko Mufwene, Linguistics
  • John Novembre, Human Genetics
  • Bruce Patterson, Field Museum
  • Catherine Pfister, Ecology and Evolution
  • Trevor Price, Ecology and Evolution
  • Victoria Prince, Organismal Biology and Anatomy
  • Stephen Pruett-Jones, Ecology and Evolution
  • Clifton Ragsdale, Neurobiology
  • Richard Ree, Field Museum
  • Olivier Rieppel, Field Museum
  • Callum Ross, Organismal Biology and Anatomy
  • Rachel Santymire, Lincoln Park Zoo
  • Urs Schmidt-Ott, Organismal Biology and Anatomy
  • Paul Sereno, Organismal Biology and Anatomy
  • Neil Shubin, Organismal Biology and Anatomy
  • Petra Sierwald, Field Museum
  • Graham Slater, Geophysical Sciences
  • Douglas Stotz, Field Museum
  • Russell Tuttle, Anthropology
  • Janet Voight, Field Museum
  • Mark Webster, Geophysical Sciences
  • Mark Westneat, Organismal Biology and Anatomy
  • John Timothy Wootton, Ecology and Evolution
  • Chung I Wu, Ecology and Evolution


  • John Bolt, Field Museum
  • Jerry Coyne, Ecology and Evolution
  • James Hopson, Organismal Biology and Anatomy
  • Michael LaBarbera, Organismal Biology and Anatomy
  • Wen-Hsiung Li, Ecology and Evolution
  • R. Eric Lombard, Organismal Biology and Anatomy
  • Thomas Nagylaki, Ecology and Evolution
  • Janice B. Spofford, Ecology and Evolution
  • Margaret Thayer, Field Museum
  • Harold Voris, Field Museum
  • William Wimsatt, Philosophy

The Committee on Evolutionary Biology (CEB) provides students with the opportunity for interdisciplinary study of all aspects of evolutionary biology. The committee consists of faculty members with primary appointments in departments in all four graduate divisions within the university and of associated faculty from institutions in the Chicago area, such as Argonne National Laboratory, Lincoln Park Zoo, Chicago Botanic Garden, the Marine Biological Laboratory, Morton Arboretum, and the Field Museum. The diversity of research interests represented by the collective expertise of the committee faculty contributes to its strong national and international reputation as a graduate training program.

Students in the committee have ready access to facilities at the associated institutions, including the more than 1,100 animals representing over 200 species at Lincoln Park Zoo, more than 17 million specimens in the Field Museum collections in botany, zoology, and paleontology, and libraries at the Field Museum. Various facilities for the study of molecular evolution and phylogenetic analysis are available to committee students, as are several student computer centers, an on-campus greenhouse, and digital equipment for off-site research.

In the Chicago area, committee students have access to the rich and diverse resources available at the Chicago Botanic Garden, Argonne National Laboratory, the Shedd Aquarium, the Morton Arboretum, and the many parks and lands managed by the local forest preserve and park districts.

The University of Chicago is a member of the Organization for Tropical Studies. Doctoral students in the committee have taken courses in tropical ecology and conducted research in Costa Rica through this affiliation. Recent evolutionary biology students have also conducted domestic research at a variety of field sites, including the Southwest Research Station of the American Museum of Natural History, Sierra Nevada Aquatic Research Laboratory, Kellogg Biological Station, the Marine Biological Laboratory at Woods Hole, and Friday Harbor Marine Laboratory.  International research is conducted on every continent.

Program of Study

Most students in the Committee on Evolutionary Biology complete their Ph.D. program in about five and a half  years.

The first and second years consist largely of course work and individual reading and research courses, aiming toward successful defense of a dissertation research proposal by the end of the Spring Quarter in the second year of study.

First year

Entering students are expected to have received a broad undergraduate training in biology and a good background in related quantitative subjects, such as chemistry, statistics and calculus. Students who are admitted with gaps in these areas may be required to remedy their deficiencies by taking appropriate courses during their first two years in the graduate program. The committee maintains a student advisory committee, which meets three times a year with each of the first and second year students to advise them on courses available, arbitrate on which courses meet the committee’s course distribution requirements, and otherwise help students keep on track towards Ph.D. candidacy.

Second year

Second year students continue to meet with the student advisory committee until they pass their preliminary examination/dissertation proposal hearing. The first part of the second year may be taken up mostly with course work, supplemented more heavily by reading and research courses.

Reading and research requirements

CEB courses have been divided into six broad areas. Students must successfully complete a course in five of the six areas to be recommended for Ph.D. candidacy. The primary aim is that the student acquires considerable breadth in evolutionary biology; this breadth and the interdisciplinary research it permits should be the distinguishing feature of students working in the committee. In the first two years of study students generally enroll in three courses per quarter. This can be a combination of lecture, seminar, research, and reading formats.

Division of the Biological Sciences teaching assistant requirement program

During their tenure in the doctoral program, students are required to register for two evaluated teaching assistantships in two approved courses.

Dissertation proposal hearing and admission to Ph.D. candidacy

Students should select an advisor no later than Autumn Quarter of their second year. This advisor normally will become the chair of the student's dissertation proposal committee. The committee for the dissertation proposal hearing will be formed by the student and her/his advisor, subject to approval by the CEB Chair, when the student asks the CEB Chair in writing to approve her/his request to appoint the exam committee and hold the proposal hearing.

CEB students must present and defend their dissertation proposal, followed by an oral examination by a faculty committee on general issues in evolutionary biology. Students are expected to successfully defend their dissertation proposal by the end of the Spring Quarter of their second year in the Ph.D. program. After successfully defending their dissertation proposal, students may be recommended for candidacy for the Ph.D. by the CEB Chair.

Ph.D. dissertation

Upon successful completion of the dissertation proposal hearing and admission into candidacy for the Ph.D., students work on their dissertation projects in close consultation with their faculty advisor and dissertation committee. During a period of two to three years the student does primary original research, participates in seminars, discussion groups, and professional meetings and conferences, and completes the written Ph.D. dissertation. Students are expected to publish dissertation related research, and encouraged to submit a substantial part of their research for publication before Ph.D. completion. A student is expected to submit a dissertation outline and proposed timetable for dissertation completion six months before the estimated date of final defense. These plans must be approved by the advisory committee, and a copy submitted as part of the meeting report to the CEB Chair.

The Ph.D. in evolutionary biology is awarded based upon the candidate’s having:

  • Submitted a written dissertation reporting results of the student’s original research in a form suitable for publication, which must be approved by the faculty advisor and dissertation committee.
  • Successfully completed a final oral examination covering the student’s field of specialization.
  • Final approval of the dissertation by the CEB Chair and the University Dissertation Office.


We strongly advise students considering application to CEB to begin preparation of their application early in the autumn quarter, so that all materials will arrive by the December 1st deadline. The committee requires GRE General Test scores from all applicants. Foreign applicants whose first language is not English also must submit TOEFL or IELTS test scores with their application materials (

Students have the opportunity to apply for the M.S. degree while completing their work for the Ph.D. The M.S. degree is also awarded in special cases, usually in association with Ph.D. requirements for graduate students in the Committee on the Conceptual and Historical Studies of Science.

Further information also may be obtained from the program’s home at, or by sending an email to

Evolutionary Biology Courses

EVOL 30196. Cultural Evolution. 100 Units.

This course explores the nature of process of cultural evolution. After establishing a background on the characteristics of biological evolution, we consider topics in cultural evolution that explore similarities and differences between processes of biological and cultural evolution, and theoretical and conceptual innovations necessary to deal with the latter, using a variety of approaches and methodologies, including agent-based modeling, "big data" approaches, and case studies. These will include topics like: the nature of inheritance, the limits of 'memes', the role of cognitive development, the coevolution of cognition and lithic technology, the scaffolding and evolution of social support, institutions, organizations and firms, the structure of scientific communities, entrenchment and the emergence of conventions and standards, the role of technology, horizontal vs. vertical transmission, multichannel inheritance, economic markets, the nature of innovation, and the role of history.

Equivalent Course(s): PHIL 52805, SOCI 40196, CHSS 40196

EVOL 30200. Chordates: Evolution and Comparative Anatomy. 100 Units.

Chordate biology emphasizes the diversity and evolution of modern vertebrate life, drawing on a range of sources (from comparative anatomy and embryology to paleontology, biomechanics, and developmental genetics). Much of the work is lab-based, with ample opportunity to gain firsthand experience of the repeated themes of vertebrate body plans, as well as some of the extraordinary specializations manifest in living forms. The instructors, who are both actively engaged in vertebrate-centered research, take this course beyond the boundaries of standard textbook content.

Instructor(s): M. Coates     Terms Offered: Winter. L.
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence, including BIOS 20187 or BIOS 20235
Note(s): Not offered Winter 2019 - Offered Winter 2020 and every other year thereafter.
Equivalent Course(s): ORGB 30250, BIOS 22250

EVOL 30300. Key Issues in Early Vertebrate Evolution. 100 Units.

Equivalent Course(s): ORGB 31300

EVOL 31200. Data Analysis in Ecol/Evol. 100 Units.

The course provides a basic introduction to statistics for biologists. We cover experimental design and many of the potential pitfalls associated with data analysis, including pseudoreplication, multiple testing, regression effects, setting up appropriate null models, and graphical presentation. Assumptions underlying elementary tests, including non-parametric vs parametric and fixed vs random effects will be clarified. We will not cover advanced methods of analysis, beyond straightforward linear models. Students will be encouraged to analyse their own datasets using R.

Instructor(s): T. Price     Terms Offered: Autumn. will be offered in Autumn 2018
Note(s): Required text: M. Whitlock & D. Schluter, Analysis of Biological Data, 2nd edition.
Equivalent Course(s): ECEV 31200

EVOL 31201. Mammalian Evolutionary Biology. 100 Units.

This course examines mammalian evolution-the rise of living mammals from ancient fossil ancestors stretching back over 300 million years. Lectures focus on the evolutionary diversification of mammals, including anatomical structure, evolutionary adaptations, life history, and developmental patterns. Labs involve detailed comparative study of mammalian skeletons, dissection of muscular and other systems, trips to the Field Museum to study fossil collections, and studies of human anatomy at the Pritzker School of Medicine. Students will learn mammalian evolution, functional morphology, and development, and will gain hands-on experience in dissection. Taught by instructors who are active in scientific research on mammalian evolution, the course is aimed to convey new insights and the latest progress in mammalian paleontology, functional morphology, and evolution. Prerequisite(s): Second-year standing and completion of a Biological Sciences Fundamentals sequence; or GEOS 13100-13200 or GEOS 22300, or consent of instructors.

Instructor(s): Z. Luo, K. Angielczyk     Terms Offered: Autumn. L.
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence including BIOS 20187 or BIOS 20235; or GEOS 13100-13200 or GEOS 22300, or consent of instructors.
Equivalent Course(s): BIOS 23262, ORGB 31201

EVOL 31700. Macroevolution. 100 Units.

Patterns and processes of evolution above the species level, in both recent and fossil organism. A survey of the current literature, along with case studies.

Instructor(s): D. Jablonski     Terms Offered: Spring
Equivalent Course(s): GEOS 36800

EVOL 31800. Taphonomy. 100 Units.

Lecture and research course on patterns and processes of fossilization, including rates and controls of soft tissue decomposition, post mortem behavior of skeletal hard parts, concentration and burial of remains, scales of time averaging, and the net spatial and compositional fidelity of (paleo)biologic information, including trends across environments and evolutionary time. Offered alternate years.

Instructor(s): S. Kidwell
Equivalent Course(s): GEOS 36700

EVOL 31900. Topics in Paleobiology. 100 Units.

In this seminar we investigate paleobiological or multidisciplinary topics of current interest to students and faculty. Previous subjects include the origin of phyla, historical and macro-ecology, the stratigraphic record and evolutionary patterns, and climate and evolution.

Instructor(s): D. Jablonski, S. Kidwell, T. Price     Terms Offered: Autumn
Equivalent Course(s): GEOS 36900, ECEV 36900

EVOL 32245. Biomechanics: How Life Works. 100 Units.

This course will explore form and function in a diversity of organisms, using the principles of physics and evolutionary theory to understand why living things are shaped as they are and behave in such a diversity of ways. Biomechanics is at the interface of biology, physics, art, and engineering. We will study the impact of size on biological systems, address the implications of solid and fluid mechanics for organismal design, learn fundamental principles of animal locomotion, and survey biomechanical approaches. Understanding the mechanics of biological organisms can help us gain insight into their behavior, ecology and evolution.

Instructor(s): M. Westneat     Terms Offered: Spring. L. Spring.
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence. Physics useful.
Note(s): This course will include a lab and will alternate years with BIOS 22233.
Equivalent Course(s): ORGB 32245, BIOS 22245

EVOL 32400. Invertebrate Paleobiology and Evolution. 100 Units.

This course provides a detailed overview of the morphology, paleobiology, evolutionary history, and practical uses of the invertebrate and microfossil groups commonly found in the fossil record. Emphasis is placed on understanding key anatomical and ecological innovations within each group and interactions among groups responsible for producing the observed changes in diversity, dominance, and ecological community structure through evolutionary time. Labs supplement lecture material with specimen-based and practical application sections. An optional field trip offers experience in the collection of specimens and raw paleontological data. Several "Hot Topics" lectures introduce important, exciting, and often controversial aspects of current paleontological research linked to particular invertebrate groups. (L)

Instructor(s): M. Webster     Terms Offered: Autumn
Prerequisite(s): GEOS 13100 and 13200, or equivalent. Students majoring in Biological Sciences only; Completion of the general education requirement in the Biological Sciences, or consent of instructor.
Equivalent Course(s): GEOS 26300, GEOS 36300, BIOS 23261

EVOL 33700. Developmental Genetics & Evolution. 100 Units.

Equivalent Course(s): BIOS 20256

EVOL 33850. Evolution and Development. 100 Units.

The course will provide a developmental perspective on animal body plans in phylogenetic context. The course will start with a few lectures, accompanied by reading assignments. Students will be required to present a selected research topic that fits the broader goal of the course and will be asked to submit a referenced written version of it after their oral presentation. Grading will be based on their presentation (oral and written) as well as their contributions to class discussions. Prerequisite(s): Advanced undergraduates may enroll with the consent of the instructor.

Instructor(s): U. Schmidt-Ott     Terms Offered: Autumn
Prerequisite(s): Advanced undergraduates may enroll with the consent of the instructor.
Equivalent Course(s): BIOS 22306, DVBI 33850, ORGB 33850

EVOL 35300. Phylogenetic Comparative Methods. 100 Units.

This is a graduate seminar course about the uses of phylogenetic trees in evolution and ecology, emphasizing historical inference of phenotypic traits, geographic ranges, and community ecology. (This is not a course on how to infer phylogenies, or their uses in studies of molecular evolution and population genetics.) Within this scope we will focus on topics of popular interest and relevance to student research. The format of the 2-hour weekly meeting will be somewhat fluid, but I anticipate giving introductory remarks or a lecture on main topics, followed by discussion of primary literature, and opportunities to work hands-on with software (bring your own laptop). Small-group assignments will be given to develop and present short tutorials on conducting analyses of real data.

Instructor(s): R. Ree, A. Hipp

EVOL 35301. Birds of the World. 100 Units.

EVOL 35401. Reconstructing the Tree of Life: An Introduction to Phylogenetics. 100 Units.

This course is an introduction to the tree of life (phylogeny): its conceptual origins, methods for discovering its structure, and its importance in evolutionary biology and other areas of science. Topics include history and concepts, sources of data, methods of phylogenetic analysis, and the use of phylogenies to study the tempo and mode of lineage diversification, coevolution, biogeography, conservation, molecular biology, development, and epidemiology. One Saturday field trip and weekly computer labs required in addition to scheduled class time. This course is offered in alternate (odd) years.

Instructor(s): R. Ree.     Terms Offered: Autumn. L.
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence or consent of instructor
Note(s): This course is offered in alternate (odd) years.
Equivalent Course(s): BIOS 23404

EVOL 35800. Classics in Evolutionary Genetics. 100 Units.

Major classic papers in evolutionary genetics that had great impact on the development of the field are reviewed.

Instructor(s): M. Long, C-I Wu     Terms Offered: Autumn. will be offered in Autumn 2018
Equivalent Course(s): ECEV 35800

EVOL 35901. Genomic Evolution I. 100 Units.

Canalization, a unifying biological principle first enunciated by Conrad Waddington in 1942, is an idea that has had tremendous intellectual influence on developmental biology, evolutionary biology, and mathematics. In this course we will explore canalization in all three contexts through extensive reading and discussion of both the classic and modern primary literature. We intend this exploration to raise new research problems which can be evaluated for further understanding. We encourage participants to present new ideas in this area for comment and discussion.

Instructor(s): M. Long, J. Reinitz, and C-I. Wu     Terms Offered: TBD. not offered in 2018-19
Equivalent Course(s): STAT 35410, ECEV 35901

EVOL 36700. Morphometrics. 100 Units.

This graduate-level course serves as an introduction to the field of morphometrics (the analysis of organismal shape). Quantitative exploratory and confirmatory techniques involving both traditional (length-based) and geometric (landmark-based) summaries of organismal shape are introduced in a series of lectures and practical exercises. Emphasis is placed on the application of morphometric methods to issues such as (but not restricted to) quantification of intraspecific variability, interspecific differences, disparity, ontogenetic growth patterns (allometry), and phylogenetic changes in morphology. Relevant statistical and algebraic operations are explained assuming no prior background. Students are required to bring personal laptop computers, and are expected to acquire and analyze their own data sets during the course.

Instructor(s): M. Webster     Terms Offered: Winter
Equivalent Course(s): GEOS 36000

EVOL 36900. Biopsychology of Sex Differences. 100 Units.

This course will explore the biological basis of mammalian sex differences and reproductive behaviors. We will consider a variety of species, including humans. We will address the physiological, hormonal, ecological and social basis of sex differences. To get the most from this course, students should have some background in biology, preferably from taking an introductory course in biology or biological psychology.

Instructor(s): Mateo, J.     Terms Offered: Autumn
Equivalent Course(s): GNSE 30901, PSYC 31600, CHDV 30901

EVOL 36905. Topics in Conservation Paleobiology. 100 Units.

Paleobiological data from very young sedimentary records, including skeletal 'death assemblages' actively accumulating on modern land surfaces and seabeds, provide unique information on the status of present-day populations, communities, and biomes and their responses to natural and anthropogenic stress over the last few decades to millennia. This course on the emerging discipline of 'conservation paleobiology' uses weekly seminars and individual research projects to introduce how paleontologic methods, applied to modern samples, can address critical issues in the conservation and restoration of biodiversity and natural environments, including such basic questions as 'has a system changed, and if so how and when relative to suspected stressors?'. The course will include hands-on experience, either in the field or with already-collected marine benthic samples, to assess societally relevant ecological change in modern systems over time-frames beyond the reach of direct observation. Enrollment limited.

Instructor(s): S. Kidwell     Terms Offered: Winter
Prerequisite(s): Additional Notes For undergraduates: completion of GEOS 13100-13200-13300 or equivalent or completion of a 20000 level course in Palentology.
Equivalent Course(s): GEOS 36905, GEOS 26905

EVOL 38600. Apes and Human Evolution. 100 Units.

This course is a critical examination of the ways in which data on the behavior, morphology, and genetics of apes have been used to elucidate human evolution. We emphasize bipedalism, hunting, meat eating, tool behavior, food sharing, cognitive ability, language, self-awareness, and sociability. Visits to local zoos and museums, film screenings, and demonstrations with casts of fossils and skeletons required.

Instructor(s): R. Tuttle     Terms Offered: Spring
Equivalent Course(s): BIOS 13253, ANTH 21428, HIPS 21428, ANTH 38600

EVOL 40100. Grants, Publications, and Professional Issues. 100 Units.

Covers professional topics in evolutionary biology, primarily strategies in grant writing and review. Each student will work towards the submission of an application of their choice. The course meets weekly and involves extensive writing and discussion.

Instructor(s): J. Bergelson, R. Ho, M. Coates     Terms Offered: Autumn
Note(s): Open to first and second year graduate students in the Darwinian Sciences Cluster
Equivalent Course(s): ECEV 40100, ORGB 40101

EVOL 40200. Advanced Topics in Ethics for the Darwinian Sciences. 100 Units.

This course covers advanced topics in ethics relevant to senior Ph.D. students in the Darwinian Sciences. CEB students are required to successfully complete this course before being awarded the Ph.D

Instructor(s): M. Coates, P. Herendeen     Terms Offered: Winter
Prerequisite(s): Open to Ph.D. students in the Darwinian Sciences
Equivalent Course(s): ECEV 40200, ORGB 40200

EVOL 41500. Topics in Stratigraphy and Biosedimentology. 100 Units.

Seminar course using the primary literature and/or a field problem. Topic selected from the rapidly evolving fields of sequence stratigraphy, basin analysis, and animal sediment relationships.

Equivalent Course(s): GEOS 38400

EVOL 42600. Community Ecology. 100 Units.

Lectures and readings cover advanced topics in multi-species systems, and include an introduction to basic theoretical approaches.

Instructor(s): J.T. Wootton     Terms Offered: Autumn
Equivalent Course(s): ECEV 42600

EVOL 42800. Population Ecology. 100 Units.

A lecture course on the empirical and theoretical approaches to the study of natural populations, including field methodologies and quantitative approaches. Includes computer assignments.

Instructor(s): C. Pfister     Terms Offered: Winter
Equivalent Course(s): ECEV 42800

EVOL 43248. Research Methods in Behavior and Development. 100 Units.

In this graduate seminar we will discuss research design, experimental methods, statistical approaches and field techniques. Other topics will be covered depending on participant interests, such as acoustic analyses, ethogram development, event recorders, spectrophotometers, marking methods, spatial analyses and grant-writing strategies. The course is primarily designed for studies of non-human animals, although studies of human behavior, especially developmental studies, will be addressed.

Instructor(s): J. Mateo     Terms Offered: Winter
Prerequisite(s): Permission of instructor.
Note(s): Not offered 2014-15
Equivalent Course(s): CHDV 43248, CHDV 23248

EVOL 44001. Molecular Evolution I: Fundamentals and Principles. 100 Units.

The comparative analysis of DNA sequence variation has become an important tool in molecular biology, genetics, and evolutionary biology. This course covers major theories that form the foundation for understanding evolutionary forces that govern molecular variation, divergence, and genome organization. Particular attention is given to selectively neutral models of variation and evolution, and to alternative models of natural selection. The course provides practical information on accessing genome databases, searching for homologous sequences, aligning DNA and protein sequences, calculating sequence divergence, producing sequence phylogenies, and estimating evolutionary parameters.

Instructor(s): M. Kreitman     Terms Offered: Winter
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence including BIOS 20187 or BIOS 20235 and two quarters of calculus, or consent of instructor.
Equivalent Course(s): BIOS 23258, ECEV 44001

EVOL 44002. Molecular Evolution II: Genes and Genomes. 100 Units.

This course covers the knowledge and well-established evolutionary analyses of genes and genomes, as well as related areas (e.g., origination and evolution of new genes, exon-intron structure, sex-related genes, sex-determination genetic systems, transposable elements, gene regulation systems, duplication of genes and genomes, evolution of genome sizes). These topics are discussed under the processes driven by various evolutionary forces and genetic mechanisms. The analysis of these problems is conducted with the genomic context. Lectures, discussions, and experiments are combined.

Instructor(s): M. Long     Terms Offered: Spring. This course is offered in alternate (odd) years.
Prerequisite(s): BIOS 23258 or consent of instructor
Equivalent Course(s): BIOS 23259, ECEV 44002

EVOL 45500. Biogeography. 100 Units.

This course examines factors governing the distribution and abundance of animals and plants. Topics include patterns and processes in historical biogeography, island biogeography, geographical ecology, areography, and conservation biology (e.g., design and effectiveness of nature reserves).

Instructor(s): B. Patterson (odd years, lab). L., Heaney (even years, discussion)     Terms Offered: Winter
Prerequisite(s): Three quarters of a Biological Sciences Fundamentals sequence and a course in either ecology, evolution, or earth history; or consent of instructor
Equivalent Course(s): GEOG 35500, BIOS 23406, GEOG 25500, ENST 25500

EVOL 46200. Evolution and the Fossil Record. 100 Units.

This course serves as an introduction to the practical and theoretical issues involved in obtaining primary systematic data from the fossil record, and demonstrates the criticality of such data to the rigorous documentation and interpretation of evolutionary patterns. Precise topics of the seminar discussions will vary from year to year depending on relevance to student research projects and interest, but are likely to focus on issues such as (but not restricted to) practical techniques in specimen-based paleontology (including fossil preparation and photography), species delimitation (including species concepts, variability, and ecophenotypy), stratigraphic/geographic range determination (including biostratigraphic correlation), phylogeny reconstruction (including the relevance of stratigraphic data), and the importance of these topics to broader macroevolutionary issues such as diversity/disparity dynamics and the determination of evolutionary trends, rates and processes.

Equivalent Course(s): GEOS 36200

EVOL 46700. Advanced Topics in Behavioral Ecology. 100 Units.

This is a reading course covering advanced topics in behavioral ecology. The list of topics to be covered will be based in part on student interests, but may include: behavior and conservation, communication, mating systems, sexual conflict, and sperm competition. This course is designed as a graduate course, but advanced undergraduates may enroll with the permission of the instructor.

Instructor(s): S. Pruett-Jones, T. Price     Terms Offered: Winter
Equivalent Course(s): ECEV 36700

EVOL 48412. Publications, Grants, and the Academic Job Market. 100 Units.

In this graduate seminar we will discuss how to write and publish scientific articles, prepare grant applications, write CVs and job applications, and give job talks and interviews. In other words, everything you always wanted to know about being successful in academia but were afraid to ask.

Instructor(s): D. Maestripieri     Terms Offered: Autumn
Equivalent Course(s): PSYC 48412, CHDV 48412

EVOL 49401. Approaches to Teaching in The Darwinian Sciences. 100 Units.

This course will introduce different teaching philosophies and methods that address how to be an effective teacher in the Darwinian Sciences. Specifically, the course will address what skills and knowledge undergraduates need to acquire and which assignments best teach these skills. Students will prepare course syllabi, discuss different approaches to teaching, and draft a philosophy of teaching statement. The overall goal for the course is that the students think critically about the art of teaching and formulate their own thoughts on the matter to better prepare them for their own careers in teaching.

Equivalent Course(s): ORGB 49401, ECEV 49401

EVOL 49500. Teaching in Evolutionary Biology. 100 Units.

Under the supervision of University faculty, graduate students in the Evolutionary Biology may serve as teaching assistants for courses in the College and relevant Graduate Divisions. Students will be evaluated and mentored throughout the quarter by their faculty supervisor, and at the end of the quarter by enrolled students. Students must choose the instructor name from the faculty listing in the Time Schedules and register using that instructor's assigned section number.

Instructor(s): Staff
Prerequisite(s): successful fulfillment of the BSD teaching requirement and consent of instructor.

EVOL 49600. Graduate Readings in Evolutionary Biology at the Field Museum. 300.00 Units.

Directed individual reading courses supervised by CEB faculty members who are curators at the Field Museum. Students must choose the instructor name from the faculty listing in the Time Schedules and register using that instructor's assigned section number.

Instructor(s): Staff
Prerequisite(s): Consent of instructor.

EVOL 49700. Graduate Readings in Evolutionary Biology. 300.00 Units.

Directed individual reading courses in evolutionary biology supervised by CEB faculty members. Students must choose the instructor name from the faculty listing in the Time Schedules and register using that instructor's assigned section number.

Instructor(s): Staff.
Prerequisite(s): consent of instructor.

EVOL 49800. Off-Campus Grad Rsch: Evolution. 300.00 Units.

Advanced research under the direction of the faculty of the Committee on Evolutionary Biology, undertaken away from the University of Chicago campus at the Field Museum, the Chicago Zoological Park, Lincoln Park Zoo, established biological field stations under the direction of their staffs, or other locations approved by the Chair and the student's advisory committee. Students must choose the instructor name from the faculty listing in the Time Schedules and register using that instructor's assigned section number.

Instructor(s): Staff
Prerequisite(s): Consent of Instructor

EVOL 49900. Graduate Research - On Campus. 300.00 Units.

Advanced research under the direction of the faculty of the Committee on Evolutionary Biology. While any approved research problem may be pursued under this course number, special attention is called to the following research fields available in the Committee: population ecology and genetics, entomology, applied ecology, plant biology, systematics of fossil invertebrates, molluscs, problems in the systematics of arthropods, herpetology, mammalogy, ornithology, and ichthyology, theoretical biology, animal behavior, paleoecology, molecular evolution, functional morphology, evolution of development, community ecology and evolution, evolutionary paleobiology and macroevolution, and physiological ecology. Students must choose the instructor name from the faculty listing in the Time Schedules and register using that instructor's assigned section number.

Instructor(s): Staff
Prerequisite(s): Consent of Instructor

EVOL 70000. Advanced Study: Evolutionary Biology. 300.00 Units.

Advanced Study: Evolutionary Biology